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By the end of this section, you will be able to:
  • Describe the unique anatomical and morphological features of flatworms, mollusks, and annelids
  • Discuss the advantages of true body segmentation
  • Explain the key features of Platyhelminthes and their importance as parasites
  • Describe the features of animals classified in phylum Annelida
  • Describe the features of animals classified in phylum Mollusca

Flatworms (Phylum Platyhelminthes), annelids (Phylum Annelida) and mollusks (Phylum Mollusca), while extraordinarily diverse, share a number of characteristics, which suggests they are more closely related to each other than other invertebrate phyla. Unlike poriferans and cnidarians, all three of these phyla are bilateral and have 3 tissue layers. This means the presence of a head with a concentration of nervous tissues. All of these phyla are also protostomes, or "mouth first" animals.

Phylum platyhelminthes

Phylum Platyhelminthes, also known as flatworms include both predatory and parasitic species, including important parasites of humans. Flatworms have three tissue layers. The flatworms are acoelomates, so their bodies are solid between the outer surface and the cavity of the digestive system.

Physiological processes of flatworms

The free-living species of flatworms are predators or scavengers. Parasitic forms feed on the tissues of their hosts. Most flatworms, such as the planarian shown in [link] , have a gastrovascular cavity rather than a complete digestive system. In such animals, the “mouth” is also used to expel waste materials from the digestive system.Digestion is extracellular. Flatworms have an excretory system with a network of tubules throughout the body with openings to the environment and nearby flame cells, whose cilia beat to direct waste fluids concentrated in the tubules out of the body. The nervous system consists of a pair of nerve cords running the length of the body with a concentration of nerves at the anterior end of the worm, where there may also be a concentration of sensory cells. These "eyespots" are the first rudimentary vision organs, but only capable of distinguishing dark and light, as well as recognize movement.

There is neither a circulatory nor respiratory system, with gas and nutrient exchange dependent on diffusion and cell-cell junctions. This necessarily limits the thickness of the body in these organisms, constraining them to be “flat” worms.

Most flatworm species are hermaphroditic, possessing both sets of sex organs, and fertilization is typically internal. Asexual reproduction is common in some groups in which an entire organism can be regenerated from just a part of itself.

Illustration shows the digestive, nervous and excretory systems in a flat, worm-like planaria. The digestive system starts at the ventral mouth opening in the middle of the animal, and then extends to the head through the middle of the body, and toward the along the sides of the body. Many lateral branches occur along the digestive system. The nervous system has 2 cerebral ganglia at the eyes in the head, and 2 ventral nerve cords with transverse connections along the length of the body to the tail. The excretory system is arranged in 2 long mesh-like structures down each side of the body.
The planarian is a flatworm that has a gastrovascular cavity with one opening that serves as both mouth and anus. The excretory system is made up of tubules connected to excretory pores on both sides of the body. The nervous system is composed of two interconnected nerve cords running the length of the body, with cerebral ganglia and eyespots at the anterior end.

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Source:  OpenStax, Bi 101 for lbcc ilearn campus. OpenStax CNX. Nov 28, 2013 Download for free at http://legacy.cnx.org/content/col11593/1.1
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