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By the end of this section, you will be able to:
  • Explain sources of evidence for evolution
  • Define homologous and vestigial structures

The evidence for evolution is compelling and extensive. Looking at every level of organization in living systems, biologists see the signature of past and present evolution. Darwin dedicated a large portion of his book, On the Origin of Species , identifying patterns in nature that were consistent with evolution and since Darwin our understanding has become clearer and broader.

Fossils

Fossils provide solid evidence that organisms from the past are not the same as those found today; fossils show a progression of evolution. Scientists determine the age of fossils and categorize them all over the world to determine when the organisms lived relative to each other. The resulting fossil record tells the story of the past, and shows the evolution of form over millions of years ( [link] ). For example, highly detailed fossil records have been recovered for sequences of species in the evolution of whales and modern horses. The fossil record of horses in North America is especially rich and many contain transition fossils: those showing intermediate anatomy between earlier and later forms. The fossil record extends back to a dog-like ancestor some 55 million years ago that gave rise to the first horse-like species 55 to 42 million years ago in the genus Eohippus . The series of fossils tracks the change in anatomy resulting from a gradual drying trend that changed the landscape from a forested one to a prairie. Successive fossils show the evolution of teeth shapes and foot and leg anatomy to a grazing habit, with adaptations for escaping predators, for example in species of Mesohippus found from 40 to 30 million years ago. Later species showed gains in size, such as those of Hipparion , which existed from about 23 to 2 million years ago. The fossil record shows several adaptive radiations in the horse lineage, which is now much reduced to only one genus, Equus , with several species.

A series of paintings on a timeline from 55 million years ago to today showing 4 of the ancestors to the modern horse. The first in the series is Eohippus, which lived from 55 to 45 million years ago. It was a small, dog-sized, animal with 4 toes on the front feet and 3 on the back, a long tail, and a brown spotted coat. The second is Mesohippus, which lived from 40 to 30 million years ago. It was slightly larger than Eohippus with longer legs. It had 3 toes on the front and back feet. The third is Hipparion, which lived from 23 to 2 million years ago. It walked on its middle toe on each foot (now a hoof), but it still had vestiges of the remaining toes. It was much larger than Hipparion. The fourth is Przewalski’s horse, a recent but endangered horse. It is smaller and stockier than the domesticated horse with one toe (hoof) on each foot.
This illustration shows an artist’s renderings of these species derived from fossils of the evolutionary history of the horse and its ancestors. The species depicted are only four from a very diverse lineage that contains many branches, dead ends, and adaptive radiations. One of the trends, depicted here is the evolutionary tracking of a drying climate and increase in prairie versus forest habitat reflected in forms that are more adapted to grazing and predator escape through running. Przewalski's horse is one of a few living species of horse.

Anatomy and embryology

Another type of evidence for evolution is the presence of structures in organisms that share the same basic form. For example, the bones in the appendages of a human, dog, bird, and whale all share the same overall construction ( [link] ). That similarity results from their origin in the appendages of a common ancestor. Over time, evolution led to changes in the shapes and sizes of these bones in different species, but they have maintained the same overall layout, evidence of descent from a common ancestor. Scientists call these synonymous parts homologous structures. Some structures exist in organisms that have no apparent function at all, and appear to be residual parts from a past ancestor. For example, some snakes have pelvic bones despite having no legs because they descended from reptiles that did have legs. These unused structures without function are called vestigial structures . Other examples of vestigial structures are wings on flightless birds (which may have other functions), leaves on some cacti, traces of pelvic bones in whales, and the sightless eyes of cave animals.

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Source:  OpenStax, University of georgia concepts of biology. OpenStax CNX. May 28, 2013 Download for free at https://legacy.cnx.org/content/col11526/1.2
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